Fossil Invertebrates
Sponges (and associated fauna)
The sponge Arborispongia (far left) (see Rigby, 1985) is locally abundant. An unnamed sponge (middle left) is much more rare. Scallop-like and mussel-like clams and spiny productid brachiopods (middle right) are frequently found attached to them. Conulariids (far right) are usually also found in association with Arborispongia (See Babcock and Feldmann, 1986). (Click on the fossils for popup enlargements.)
Sphenothallus
Two species of the attached invertebrate Sphenothallus Hall (Hill, 1978) are also common, each species in a different environment. Species 1 (left), a solitary form, is found toward the deeper part of the bay, while Species 2 (right), a branching, colonial form, is an indicator of shallow water. Sphenothallus is known from similar aged rocks in Indiana and Glencartholm, Scotland.
Brachiopods and Bivalves
Occurrences of the inarticulate brachiopod Lingula are localized, as are the occurrences of the few kinds of articulate brachiopods (see Lutz-Garihan, 1985). Scallop-like and mussel-like bivalves, as well as Sphenothallus species 1, are found attached to many things; such as algal fronds (see McRoberts and Stanley, 1989), sponges, a sunken log, and cephalopod shells.
Cephalopods
Several species of orthoconic (Reticycloceras) and coiled (Anthracoceras) nautiloid cephalopods are abundant. While the shell minerals are usually gone, many specimens preserve traces of internal anatomy. They are indicators of open water marine influence (see Landman and Davis, 1988; Mapes, 1987).
Assorted Worms
There are many species of platyhelminth, nemertine, and annelid worms in the fauna (see Schram 1979c). (Click on the fossils to view a popup enlargment.)
Arthropods
At least one dozen species of concavicarid, phyllocarid (Dithyrocaris view 1 & view 2), paleostomatopod (Bairdops and Tyrannophontes), and eumalocostracan (Aenigmocaris) crustaceans are known (see Briggs and Rolfe, 1983; Factor and Feldmann, 1985; Hof, 1998; Jenner et al., 1998; Schram and Horner, 1978; Schram, 1979a, 1985). The sole species of horseshoe crab, Paleolimulus, is rare (see Schram, 1979b). Several species of Bear Gulch fish have crustaceans in their guts. Only one tiny trilobite specimen has been found in the Bear Gulch Limestone although they are not rare in adjacent shelly beds of the Heath Formation.
Echinoderms
Echinoderms (crinoids, echinoids, asteroids, and ophiuroids) such as Lepidasterella are rare (see Welch, 1984), highly localized, and usually poorly preserved because of the postmortem loss of aragonite from their skeletons.
References
- Babcock, L. E., and R. M. Feldmann, 1986. "Devonian and Mississippian conulariids of North America. part B. Paraconularia, Reticulaconularia, New Genus, and organisms rejected from Conulariida." Annals of Carnegie Museum 55: 411-479. (back)
- Briggs, D. E. G., and W. D. I. Rolfe, 1983. "New Concavicarida (new Order: ?Crustacea) from the Upper Devonian of Gogo, Western Australia, and the paleoecology and affinities of the group". Spec. Papers in Palaeontology 30: 249-276. (back)
- Cox, R. S. 1986. "Preliminary report on the age and palynology of the Bear Gulch Limestone (Mississippian, Montana)". Journal of Paleontology 60: 952-956.
- Factor, D., and Feldmann, R., 1985. "Systematics and paleoecology of malacostracan arthropods in the Bear Gulch Limestone (Namurian) of Central Montana. Annals of Carnegie Museum 54: 319-356. (back)
- Hill, V. 1978. Sphenothallus cf. S. angustifolius from the Mississippian Bear Gulch Limestone of Central Montana". Unpublished B.A. thesis, Princeton Univ. Geology Dept. 30 pp. + plates and appendix. (back)
- Hof, C. H. J. 1998. "Fossil Stomatopods (Crustacea : Malacostraca) and their phylogenetic impact". Journal of Natural History 32:1567-1576. (back)
- Jenner, R. A., C. H. J. Hof, and F. R. Schram. 1998. "Palaeo- and archaeostomatopods (Hoplocarida: Crustacea) from the Bear Gulch Limestone, Mississippian (Namurian), of Central Montana." Contributions to Zoology 67:155-185. (back)
- Landman, N. H., and R. A. Davis, 1988. "Jaw and crop preserved in an orthoconic nautiloid cephalopod from the Bear Gulch Limestone (Mississippian, Montana)." New Mexico Bureau of Mines and Mineral Resources mem. 44:103-107. (back)
- Lutz-Garihan, A. B. 1985. "Brachiopods from the Upper Mississippian Bear Gulch Limestone of Montana." Compte Rendu, Neuvième Congrès International de Stratigraphie et de Géologie du Carbonifère, 5: 457-467. (back)
- Mapes, R. S. 1987. "Upper Paleozoic cephalopod mandibles: frequency of occurrence, modes of preservation, and paleoecological implications". Journal of Paleontology 61: 521-538. (back)
- McRoberts, C.A., and G.D. Stanley. 1989. "A unique bivalve-algae life assemblage from the Bear Gulch Limestone (Upper Mississippian) of central Montana." Journal of Paleontology 63:578-581. (back)
- Rigby, J. K. 1985. "The sponge fauna of the Mississippian Heath Formation of Central Montana." Compte Rendu, Neuvième Congrès International de Stratigraphie et de Géologie du Carbonifère, 5: 443-456. (back)
- Schram, F. R. 1979a. "The genus Archaeocaris, and a general review of the Paleostomatopoda (Hoplocarida: Malacostraca)". Trans. San Diego Soc. Nat. Hist. 19: 57-66. (back)
- Schram, F. R. 1979b. "Limulines of the Mississippian Bear Gulch Limestone of Central Montana, USA." Trans. San Diego Soc. Nat. Hist., 19:67-74. (back)
- Schram, F. R. 1979c. "Worms of the Mississippian Bear Gulch Limestone of Central Montana, USA." Trans. San Diego Soc. Nat. Hist. 19: 107-120. (back)
- Schram, F. R. 1985. "The Bear Gulch crustaceans and their bearing on Late Paleozoic diversity and Permo-Triassic evolution of Malacostraca." Compte Rendu, Neuvième Congrès International de Stratigraphie et de Géologie du Carbonifère, 5:468-472. (back)
- Schram, F. R., and J. R. Horner, 1978. "Crustacea of the Mississippian Bear Gulch Limestone of Central Montana", Journal of Paleontology 52: 394-406. (back)
- Welch, J. R. 1984. "The asteroid, Lepidasterella montanensis n. sp., from the Upper Mississippian Bear Gulch Limestone of Montana." Journal of Paleontology 58: 843-851. (back)
