Ph.D. (1993) in Marine Science, College of William and Mary
M.S. (1988) in Biology, Adelphi University
B.S. (1984) in Biology, Washington College
Professor of Biology, Saint Joseph's University
Similarity of morphological composition and developmental patterning in paired fins of the elephant shark. Riley, Cloutier, and Grogan 2017. www.nature.com. Scientific Reports 7: 9985 | DOI:10.1038/s41598-017-10538-0
2016 Tengelmann Award
I take an integrative approach (fossil and extant) to the study of the origin and relationships of chondrichthyan fishes, and early vertebrates in general. Emphasis is placed on the functional and developmental morphology of the cranium, suspensorium, mineralized tissues and the immune and vascular systems. Studies in progress involve ontogenetic and phylogenetic differentiation of the cranium, the development and mineralization of skeletal tissues, the description of new Bear Gulch fishes, and phylogenetic and ecomorphological analyses of these Paleozoic fishes.